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Meanwhile, Carl Linnaeus (1707–1778), the Swedish physician, botanist, and zoologist, modified the established taxonomic bases of binomial nomenclature for fauna and flora, and was a pioneer researcher in biologically defining human race . In Systema Naturae (1767), he labeled five  " varieties "   of human species. Each one was described as possessing the following physiognomic characteristics "varying by culture and place" :